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LSI PhD Programme in Complex Living Systems

LSI PhD Programme

LSI PhD Programme

LSI PhD Programme

LSI PhD Programme

Modern scientific innovation involves the integration of diverse disciplines and the bridging of pure and application-oriented research. At the LSI we are training a new generation of scientists who are equipped to think and experiment beyond traditional boundaries using cutting-edge technologies and approaches.

You can apply to our PhD Programme via three routes: (i) direct application for an LSI-funded PhD studentship (covers fees, stipend, and training budget); (ii) application for a studentship funded by another source (e.g., UKRI, China Scholarship Council, national agencies) with an LSI supervisor; (iii) self-funding with an LSI supervisor.

Hear from our students

What we offer

All students are enrolled in the integrated LSI programme regardless of primary discipline or funding source. You will benefit from the broad range of opportunities and activities offered by the programme, including:

  • Leading-edge research technologies and approaches, from quantum biology to brain function
  • State-of-the-art laboratories and facilities in the recently built LSI
  • Opportunity to formulate and modify your own project
  • Cohort learning in interdisciplinary training modules and workshops
  • Supervisors from different disciplines
  • Advisory thesis committee to support you throughout the PhD
  • Diverse and inclusive community of over 60 PhD students
  • Peer-to-peer PhD student presentation and discussion sessions
  • Future career preparation

Exeter is one of the fastest growing cities in the UK.  It is situated in the South West of England which offers unmatched sports and leisure opportunities through several national parks and some of the best beaches in Europe.


Who we’re looking for

We aim to recruit the best and most imaginative students from across the range of disciplines represented in LSI – Physics, Mathematics, Computational Science, Engineering, Biology and Medicine.

You will be motivated by scientific curiosity and be able and willing to “think outside of the box”. You should have a first or upper-second-class degree (or equivalent). Masters training can be an advantage but is not essential. However, you should have sufficient experience to demonstrate your capability for research work and aptitude for critical thinking. 


Project titles for LSI-funded PhD studentships

(starting Sept 2024)

  • “Mathematical modelling of pattern formation mediated by signalling filopodia”;  Supervisors: Kyle Wedgwood (Maths), Steffen Scholpp (Biosciences)

  • "Modelling shape generation by the basement membrane"; Supervisors: Stefan Harmansa (Bio) and David Richards (Physics)

  • "Novel high-throughput microfluidic technologies for the screening of bacteriophages eradicating bacterial biofilms"; Supervisors: Fabrice Gielen (Physics) Remy Chait (Bio)

  • "Peptide magnetosensors: new quantum mechanisms to explain and engineer magnetic field sensitivity in biological systems"; Supervisors: Daniel Kattnig (Physics) and Jonathan Phillips (Bio)

  • "Structural dynamics of genome editing enzymes: A combined mathematical and experimental approach"; Supervisors: Jonathan Phillips (Bio) and Marc Goodfellow (Maths)

  • "The role of fluid flow and luck in the coevolution of microbial interactions in the vast ocean"; Supervisors: Wolfram Moebius (Physics), Adam Monier (Bio), Daniel Katting (Physics)

  • "Understanding stress-induced changes in RNA processing"; Supervisors: Soojin Ryu (CBS) and Steven West (Bio)

  • "Unlocking Nature's Compass: Probing the Quantum Secrets of Avian Magnetoreception with Single Molecule Sensors"; Supervisors: Frank Vollmer (Physics) and Daniel Kattnig (Physics)

  • "Unlocking the molecular mechanisms governing stem cell dormancy"; Supervisors: Bertram Daum (Bio) and Austin Smith (Bio)



Applications for LSI-funded PhD studentships are open now.

The deadline will be midnight 8th January 2024

Apply now

For other studentships deadlines see details on individual research group pages.

Other funded studentships deadlines vary. Please see the details and links on the group pages.

For self-funded applicants, please contact directly your preferred supervisor and also the LSI Director of Postgraduate Research, Professor Steve West (

In your message, please state your motivation for carrying out Ph.D. research, why you are applying to our interdisciplinary programme, why you have chosen your selected supervisor/project, and your source of funding for fees, living costs and research training. Self-funded applicants will be subject to an equivalent selection process as applicants for funded positions.

If you have any inquiries, please contact:

Director of Postgraduate Research (LSI), Steven West:


LSI PhD Studentships - funded by different sources


Funding body: BBSRC SWBio DTP and Syngenta

Title: Time-resolved kinetic and structural understanding of essential enzymes to develop novel herbicides

Supervisors: Nicholas Harmer (Exeter), Jim Spencer (Bristol), Christian Noble (Syngenta)

Application Deadline: Monday 4th December 2023   Application website 


Funding BBSRC, SWBio DTP

Title: Studying early human embryo development using stem cell models

Supervisors: Ge Guo (Exeter), Fabrice Gielen (Exeter), Rosalind John

Project information 

Application Deadline: Monday 4th December 2023  Application website


Funding BBSRC, SWBio DTP

Title: Dynamic Protein Design

Supervisors: Jonathan Phillips (Exeter) and Marc Goodfellow (Exeter)

Application Deadline: Monday 4th December 2023   Application website


Funding BBSRC, SWBio DTP

Title: Characterizing in vivo cell cell interaction in stress system

Supervisors: Soojin Ryu (Exeter-LSI ), Joel Tabak (Exeter-Math), Jamie Walker (Exeter-Math)

Application deadline: Monday 4th December   Application website



Funding BBSRC, SWBio DTP

Title: Spindle orientation in the developing fly embryo: a joint mathematical-experimental approach

Supervisors: David Richards (Exeter) and James Wakefield (Exeter)

Application Deadline: Monday 4th December 2023   Application website 


Funding body: NERC GW4+ DTP PhD studentship 

Title: Comparative analysis and modelling of cilia motility in a major disease-causing parasite, Supervisors

Supervisors: Kirsty Wan, University of Exeter, Aidan Emery, Natural History Museum, Bonnie Webster, Natural History Museum, Adam Cieplinski, Natural History Museum

Application Deadline: Tuesday 9th January 2024   Application website